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Expendable, queenless colonies are being considered seriously as a substitute for normal colonies when bees are needed in locations where pesticide exposure is too severe to risk loss or damage to normal colonies. VII. Swarming and Swarm Migration Swarming and supersedure behavior apparently are related to the production and distribution of queen phero mones. Butler (1960b) found that superseded and "swarm" queens from meagerly-populated colonies had approximately 25% as much queen substance as mated, laying queens.
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